| Haroldson, Ternent, Gunther, Schwartz | Mark A., Mark A., Kerry A., Charles C. | Grizzly bear denning chronology and movements in the Greater Yellowstone Ecosystem | 2002 | Ursus 13:29-37. |
Excerpts from the Abstract:
Den entry for females began during the fourth week in September, with 90% denned by the fourth week of November. Earliest den entry for males occuered durring the second week of October, with 90% denned by the second week of December.
Earliest den emergence for males occurred durring the first week of February, with 90% of males out of dens by the forth week of April. Earliest den emergence for females occured during the third week of March; by the first week of may, 90% of female had emerged.
Known pregnant females tend to den at high elevation and, following emengence, remained at higher elevation until late May. Females with cubs remained relatively close (<3km) to den sites until the last two weeks in May.